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In Model I the DNA would bend toward the protein [as observed with the catabolite activator protein (CAP) from Escherichia coli (20) and the eukaryotic nucleosome core particle (21)].
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A discrepancy does occur for Sm and Eu but this is due to low-lying excited states with different J from the ground states which, because of their close proximity to the ground state in energy, are also significantly populated and cause p to shift from its ground state value.
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